The Genesect Army comes to a mountainous, snowy area that used to be their home. However, three hundred million years have passed, and it is no longer habitable. Suddenly, an avalanche occurs, but Mewtwo arrives and manages to save four of the five Genesect. Mewtwo peers into the mind of the Douse Drive Genesect and learns what they are looking for: their home. The Red Genesect, whom Mewtwo was unable to save, then appears and attacks Mewtwo, and the Genesect Army flies off. Later, they see the skyline of a large city and mistake it for their home, leading them to travel there.
Ash, Cilan, and Iris arrive in New Tork City, where their guide, Eric, allows them to visit Pokémon Hills, a nature park, before it opens to the public. While there, Ash finds and quickly befriends the Douse Drive Genesect. The Red Genesect and Shock Drive Genesect appear and think that Ash is hurting the Douse Drive Genesect, causing them to attack him; the Red Genesect forces the Douse Drive Genesect to attack Ash as well. Mewtwo shows up and protects Ash and his friends from the Genesect, causing them to leave.
That night, the Genesect begin taking over the center of Pokémon Hills, driving out the other Pokémon and building a giant cocoon. The cocoon threatens the city's power supply, so Ash, Iris, and Eric go underground to try to fix the problem. Unsuccessful, they then enter the center of Pokémon Hills from below, where the Douse Drive Genesect apologizes for attacking Ash earlier, making Ash think that it is unable to disobey the Red Genesect. Iris realizes that the cocoon is the Genesect Army's nest and that the Genesect are trying to turn Pokémon Hills into their new home. A Feraligatr native to Pokémon Hills starts attacking the nest and the Red Genesect but is knocked off a ledge.